Unidentified spirits begin appearing in Karakura Town. A mysterious Soul Reaper who calls herself Senna appears and begins destroying the spirits. Ichigo Kurosaki and Rukia Kuchiki confront her about the situation, but she refuses to answer any questions and leaves. Ichigo follows her while Rukia searches for answers. Ichigo meets up with Toshiro Hitsugaya and Rangiku Matsumoto at the Urahara Shop, who have been sent to earth to investigate why the real world is suddenly appearing as a reflection in the sky in Soul Society.
Kisuke Urahara explains that a dimension between Soul Society and the real world, the Valley of Screams, has expanded to the point that it is connecting the two worlds. The spirits seen earlier are "Blanks", souls without memories that were lost in the space between the two worlds and that comprise the Valley of Screams. The memories of the Blanks combine to form a single being, the "Memory Rosary." Upon further investigation, it is revealed that Senna is this being.
The Gotei 13 are sent to try to deal with the problem of the expanding valley, however the "Dark Ones", a group of exiles from the Soul Society seeking revenge for the past, appear and capture Senna. They take her to the Valley of Screams and attach her to a device powered by the Blanks that will cause the Valley to collapse, resulting in a collision between the real world and the Soul Society that would destroy both worlds.
Ichigo travels to the Valley to rescue Senna while Rukia goes to get reinforcements from Soul Society. However, General Yamamoto wants to destroy the Valley before it collapses and refuses to order a rescue mission with only an hour left. In the Valley, Ichigo is overpowered by the large number of blanks, but he is saved by the arrival of his many friends from the Soul Society. While the group battles the Dark Ones and Blanks, Ichigo battles and eventually defeats their leader, Ganryu.
The process of the collapse has gone too far to completely stop. Senna sacrifices herself to push the two worlds back apart to save Ichigo. Senna asks Ichigo to take her to the graveyard so she can see her name on her gravestone. Though the stone is blank, Ichigo lies and tells her that her name is on it. Believing him, she expresses contentment that she once had a life of her own before fading away. Rukia notes that once the power of the Blanks fades away, so will all memories of Senna. After the credits, Ichigo is walking on the bridge where the portal was and sees a red ribbon, the one that he had bought Senna, floating down from the sky then sees a girl who looks just like her running past him, he smiles and continues on with his walk.
h-roh berwarna putih yang berwujud aneh, tidak dikenali oleh siapa pun, mulai bermunculan di Kota Karakura. Seorang gadis Shinigami yang menyebut dirinya Senna muncul bersamaan dengan kemunculan mereka dan menghabisi roh-roh tersebut dengan zanpakutou-nya. Kurosaki Ichigo dan Kuchiki Rukia yang bertanya-tanya melihat situasi tersebut segera menghadap Senna, namun gadis itu tidak menjawab dan melarikan diri dari mereka.
Di sisi lain, di langit Soul Society mulai terlihat refleksi dari dunia manusia, pertanda bahwa mulai terbukanya dimensi yang menghubungkan Soul Society dan dunia manusia, Valley of Scream. Roh-roh putih yang berkeliaran di Kota Karakura ditengarai sebagai salah satu penyebabnya, dan terdapat sebuah "Memory Rosary" yang harus digunakan untuk mencegah terjadinya pencampuran antara dunia manusia dan Soul Society.
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